Title Page

Contents

ABSTRACT 7

1. Introduction 8

1.1. Introduction to the family Nolidae 8

1.2. The historical review of the Korean Nolidae 9

1.3. Morphology 11

Morphology of the adult 11

Morphology of genitalia 16

Immature stage 18

2. Materials and Methods 20

2.1. Methods 20

2.2. Preparation of genitalia and abdomen 21

2.3. Preparation of wing venation 22

2.4. Abbreviations 23

3. Systematics 24

3.1. Checklist of Korean Nolidae 24

3.2. Taxonomic accounts 29

Subfamily Risobinae Mell, 1943 29

Subfamily Eligminae Mell, 1943 32

Subfamily Westermanniinae Hampson, 1918 40

Subfamily Nolinae Hampson, 1894 42

Subfamily Chloephorinae Stainton, 1859 121

3.3. Phylogeny 168

Characters used in cladistic analysis 168

Cladistic analysis 174

DNA barcoding for Meganola bryophilalis (Staudinger) species-complex 175

4. Conclusions 176

5. References 178

6. Appendix 183

6.1. Type specimen of Nola kanshireiensis (Wileman & South, 1916), and N. yoshinensis (Wileman & West, 1929) 183

6.2. DNA barcodes used in this study 184

국문요약 192

Legend of Plates 193

Adult plates 1-22. 194

Male genitalia plates 1-25. 238

Female genitalia plates 1-18. 288

Venation plates 1-7. 322

List of the author publications 342

Book 342

Journal 342

Table 1. Data matrix for each genus of Nolidae (outgroup: Arctia; ingroup: 28 genera; in total 62 characters) 172

Text figure 1. Head part of the family Nolidae. 11

Text figure 2. Sound producing structure, tymbal organ of Pseudoips prasinana (Linnaeus). 12

Text figure 3. Tympanal structure of Nolidae (Macrochthonia fervens Butler). 13

Text figure 4. Representative wing venation of Nolidae (Meganola subgigas Inoue). 14

Text figure 5. Bar-shaped retinaculum of male Nolidae. 15

Text figure 6. Representative male genitalia of Nolidae (Manoba banghaasi (West) & Meganola ohsunghwni László, Ronkay & Ronkay). 16

Text figure 7. Representative female genitalia of Nolidae (Meganola subgigas Inoue). 17

Text figure 8. Larva of Meganola ohsunghwani László, Ronkay & Ronkay. 18

Text figure 9. Pupa of Blenina senex (Butler) 19

Text figure 10. Parsimony tree from the unordered option with characters of 29 genera of the family Nolidae with an outgroup from Korea using PAUP 4.0a169 (outgroup＝Arctia Schrank) 174

Text figure 11. Neighbor-Joining tree of selected nolid moths 175