Title Page
Contents
요약 7
I. Introduction 11
II. Historical background 16
1. Botanical study 16
2. Study of ferns 18
III. Materials and Methods 24
1. Field survey 24
2. Taxon sampling 27
3. Classification systems 27
4. Literatures 28
IV. Results and Discussion 30
1. Characteristics of ferns and lycophytes in Cambodia 30
1) List of ferns 30
2) Distribution by province 36
3) Distribution by altitude 38
4) Distribution by ecology 41
5) Phytogeographical patterns 44
2. New Taxa 48
1) xCephalodictyum thambangensis B.-Y. Sun & I.C. Hwang, hybrid nov. 48
2) Lindsaea kohkongensis B.-Y. Sun & I. C. Hwang sp.nov. 54
3. Taxonomic treatment 61
1. Lycopodiaceae P. Beauv. ex Mirb., Hist. Nat. Vég. Classés par Familles 4: 293. 1802. 66
2. Selaginellaceae Willk. Anleit, Stud. Bot. 2: 163. 1854. 75
3. Isoetaceae Dumort., Anal. Fam. Pl. 67. 1829. 106
4. Psilotaceae J. W. Griff. & Henfr., Microgr. Dict.540. 1855. 108
5. Equisetaceae Michx. ex DC., Essai Propr. Méd. Pl. 49. 1804. 110
6. Ophioglossaceae Martinov, Tekhno-Bot. Slovar. 438. 1820. 112
7. Marattiaceae Kaulf. Enum. Filic. 31. 1824. 118
8. Osmundaceae Martinov, Tekhno-Bot. Slovar. 445. 1820. 122
9. Hymenophyllaceae Mart., Consp. Regn. Veg. 3. 1835. 124
10. Gleicheniaceae C. Presl, Reliq. Haenk. 1: 70. 1825. 152
11. Dipteridaceae Seward & E. Dale, Phil. Trans. R. Soc. B 194: 487, 499, 502. 1901. 164
12. Lygodiaceae M. Roem., Handb. Allg. Bot. 3: 520. 1840. 166
13. Schizaeaceae Kaulf., Wesen Farrenkr. 119. 1827. 175
14. Marsileaceae Mirb. in Lam. & Mirb. Hist. Nat. Vég. 5: 126. 1802. 178
15. Salviniaceae Martinov, Tekhno-Bot. Slovar. 559. 1820. 181
16. Plagiogyriaceae Bower, Ann. Bot. (London) 40: 484. 1926. 185
17. Cibotiaceae Korall, Taxon 55(3): 712. 2006. 187
18. Cyatheaceae Kaulf., Wesen Farrenkr. 119. 1827. 189
19. Lindsaeaceae C. Presl ex M.R. Schomb., Reis. Br.-Guiana 3: 883, 1048. 1848. 199
20. Pteridaceae E.D.M. Kirchn., Schul.-Bot. 109. 1831. 219
21. Dennstaedtiaceae Lotsy, Vortr. Bot. Stammesgesch. 2: 655. 1909. 275
22. Aspleniaceae Newman, Hist. Brit. Ferns 6. 1840. 295
23. Blechnaceae Newman, Hist. Brit. Ferns, ed. 2: 8. 1844. 318
24. Athyriaceae Alston, Taxon 5: 25. 1956. 325
25. Thelypteridaceae Pic. Senn., Webbia 24: 709. 1970. 335
26. Lomariopsidaceae Alston, Taxon 5: 25. 1956. 374
27. Dryopteridaceae Herter, Revista Sudamer. Bot. 9: 15. 1949. 378
28. Tectariaceae Panigrahi, J. Orissa Bot. Soc. 8: 41. 1986. 416
29. Oleandraceae Ching ex Pic. Serm., Webbia 20(2): 745. 1965. 435
30. Nephrolepidaceae Pic. Serm., Webbia 29(1): 8-11. 1974. 441
31. Davalliaceae M.R. Schomb., Reis. Br.-Guiana 3: 883. 1848. 451
32. Polypodiaceae J. Presl & C. Presl, Delic. Prag.:159. 1822. 467
4. Conclusion 558
Abstract 560
References 565
Appendix 1. List of ferns and lycophytes in Cambodia 576
Table 1. Survey dates and regions 24
Table 2. Summary of the checklist of ferns and lycophytes in Cambodia 31
Table 3. Taxa found only in the literatures 34
Table 4. List of species showing restricted distribution in particular area 37
Table 5. Division of ecology type 42
Table 6. Comparision of characters in Cephalomanes javanicum, × Cephalodictyum thmabangensis and Abrodictyum obscurum 51
Table 7. Comparision of characters in L. ensifolia, L. kohkongensis and L. heterophylla 57
Fig 1. Research areas in the literatures 23
Fig 2. Survey areas in Cambodia 26
Fig 3. Number of species distribution at each region 37
Fig 4. Distribution of species and its maximum altitude 39
Fig 5. Distribution of species and its minimum altitude 40
Fig 6. Number of taxa by altitudinal belt 41
Fig 7. Distribution rate of ecology type 44
Fig 8. A comparition of the species of ferns in Cambodia and other neighboring countries 45
Fig 9. Distribution rate of ferns in Cambodia on the Asiatic tropics 46
Fig 10. Distribution of the Main Taxanomic Group of the Cambodian Ferns in the Neighboring Countries 48
Fig 11. Comparision of plants in 1-Abrodictyum obscurum, 2-Cephalomanes javanicum and 3-× Cephalodictyum thmabangensis 52
Fig 12. Comparision of Cells in 1-Cephalomanes javanicum, 2-×Cephalodictyum thmabangensis and 3-Abrodictyum obscurum 52
Fig 13. ×Cephalodictyum thmabangensis 53
Fig 14. Habitate and Plants of L. kohkongensis 58
Fig 15. Comparision of Plant in 1, 2-L. ensifolia, 3-L. heterophylla and 4-L. kohkongensis 59
Fig 16. Lindsaea kohkongensis 60